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Admissions Information

Application Deadlines

Degree Requirements


Financial Assistance







Assistant Dean

Thompson, Katerina, Adjunct Member

Associate Director

Brooks, Michelle,

Distinguished University Professor

Colwell, Rita R., Full Member

Gantt, Elisabeth, Full Member


Allewell, Norma M., Full Member

Andrews, Norma, Full Member

Carleton, Karen, Full Member

Carr, Catherine E., Full Member

Chang, Caren, Full Member

Colombini, Marco, Full Member

Cooke, Todd J., Full Member

Cummings, Michael P., Full Member

Delwiche, Charles Francis, Full Member

DeStefano, Jeffrey J., Full Member

Dinman, Jonathan D., Full Member

Fagan, William F., Full Member

Feijo, Jose, Full Member

Hannenhalli, Sridhar,

Hutcheson, Steven W., Full Member

Jeffery, William Richard, Full Member

Kanold, Patrick, Full Member

Kocher, Thomas D., Full Member

Lips, Karen, Full Member

Machado, Carlos, Full Member

Mather, Ian H., Full Member

Mitter, Charles, Full Member

Mosser, David M., Full Member

Palmer, Margaret, Full Member

Payne, Richard, Full Member

Porter, Tom E., Full Member

Quinlan, Elizabeth, Full Member

Simon, Jonathan, Full Member

Simon, Anne E., Full Member

St.Leger, Raymond, Full Member

Stein, Daniel C., Full Member

Sukharev, Sergei I., Full Member

Via, Sara, Full Member

Wilkinson, Gerald S., Full Member

Wolniak, Stephen M., Full Member

Associate Professor

Ades, Ibrahim Z., Full Member

Araneda, Ricardo, Full Member

Belov, Georgiy, Full Member

Bely, Alexandra E., Full Member

Butts, Daniel, Full Member

Cao, Kan,

El-Sayed, Najib, Full Member

Haag, Eric S., Full Member

Hawthorne, David, Full Member

Kahn, Jason D., Full Member

Lamp, William, Full Member

Liu, Zhongchi, Full Member

McIver, Kevin, Full Member

Mount, Stephen M., Full Member

Neel, Maile, Full Member

Pick, Leslie, Full Member

Shultz, Jeffrey, Full Member

Singer, Joshua, Full Member

Song, Wenxia, Full Member

Stewart, Richard C., Full Member

Straney, David C., Full Member

Swenson, Nathan,

Wang, Jian, Full Member

Assistant Professor

Briken, Volker, Full Member

Dwyer, Daniel,

Fisher, Heidi, Full Member

Fritz, Megan, Non-Member

Gaudry, Quentin, Full Member

Gruner, Daniel, Full Member

Hamby, Kelly, Full Member

Johnson, Philip,

Jose, Antony,

Juntti, Scott, Full Member

Lee, Vincent, Full Member

Roy, Sougata,

Song, Jiuzhou, Full Member

Speer, Colenso, Full Member

Taneyhill, Lisa, Full Member

Adjunct Professor

Braun, Michael, Adjunct Member

Collins, Allen, Adjunct Member

DiMichele, William A., Adjunct Member

Evans, Jay, Adjunct Member

Fleischer, Robert, Non-Member

Freed, Eric, Adjunct Member

Green, Kim, Adjunct Member

Hines, Anson H., Adjunct Member

Kelley, Matthew W., Adjunct Member

Labandeira, Conrad, Special Member

Leimgruber, Peter, Non-Member

Marino, John, Full Member

Marra, Peter, Adjunct Member

McBride, Alison, Adjunct Member

Moss, Bernard, Adjunct Member

Moult, John, Full Member

Parr, Cynthia, Adjunct Member

Rosenthal, Benjamin M., Adjunct Member

Stoltzfus, Arlin, Adjunct Member

Sues, Hans, Non-Member

Wing, Scott, Adjunct Member

Zimmer, Elizabeth, Adjunct Member

Affiliate Professor

Bentley, William E., Full Member

Chaverri, Priscila, Full Member

Culver, James N., Full Member

Darden, Lindley, Full Member

Dooling, Robert J., Full Member

Hamza, Iqbal, Full Member

Herzberg, Osnat, Full Member

Kelman, Zvi, Full Member

Mariuzza, Roy, Full Member

Murphy, Angus S., Full Member

Nelson, Daniel, Full Member

Nuss, Donald, Full Member

Pal, Utpal, Full Member

Pop, Mihai, Full Member

Wu, Louisa, Full Member

Xiao, Shunyuan, Full Member

Zhu, Xiaoping, Full Member

Affiliate Associate Professor

Corrada Bravo, Hector, Full Member

Hsieh, Adam H., Full Member

Muro, Sylvia, Full Member

Zhang, Yanjin, Full Member

Affiliate Assistant Professor

Biswas, Debabrata, Full Member

Jay, Steven M., Full Member

Jewell, Christopher M., Full Member

Kim, Byung-Eun, Full Member

Micallef, Shirley, Full Member

Telugu, Bhanu P., Full Member

Professor Emerita

Cohen, Avis H., Full Member

Sze, Heven, Full Member

Professor Emeritus

Gill, Douglas E., Full Member

Highton, Richard, Full Member

Inouye, David W., Full Member

Popper, Arthur N., Full Member

More BISI Faculty Information

Biological Sciences (BISI)   

Program Title and Classification

Biological Sciences

Graduate Degree Program

College: Computer, Mathematical, and Natural Sciences


The Biological Sciences (BISI) Graduate Program offers a wide range of training opportunities for students interested in pursuing doctoral level research in exciting, diverse areas across the biological sciences. BISI is an interdepartmental umbrella program that consists of four Areas of Concentration:

Behavior, Ecology, Evolution, and Systematics (BEES)

Computational Biology, Bioinformatics, and Genomics (CBBG)

Molecular and Cell Biology (MOCB)

Physiological Systems (PSYS)

Graduate students in BISI are associated with one of these areas, but their interests often span multiple concentrations, which allows the development of  innovative research projects across traditional disciplinary boundaries. Descriptions of each Concentration Area, faculty research interests, and more detailed programmatic information are available at

Although the BISI Program is administered within the College of Computer, Mathematical, and Natural Sciences, it involves distinguished graduate faculty from multiple departments and Colleges at the University of Maryland as well as outstanding adjunct faculty from nearby research institutions. Students may have opportunities to work with participating scientists from - as examples - the National Institutes of Health; Smithsonian Institution, including the National Museum of Natural History, the National Zoo, and Molecular Systematics Laboratory; the Food and Drug Administration; and the U.S. Department of Agriculture. BISI students have an incomparable wealth of potential research options and collaborations that extend from Maryland’s College Park campus throughout the Washington D.C. metropolitan area.

Admissions Information

General Requirements:

Program-Specific Requirements:

  • Letters of Recommendation (3)
  • Graduate Record Examination (GRE)
  • CV/Resume
  • Supplementary Application
  • Publications/Presentations

All students applying to the Biological Sciences Graduate Program must have a Bachelor’s degree from a recognized undergraduate institution. Applicants are expected to have a strong academic record, including coursework in advanced areas of biology as well as at least one year of calculus, general chemistry, organic chemistry, and physics. Able students with deficiencies in a particular area may be admitted and the deficiency corrected after enrollment.  Students are asked to identify a concentration area and research interests on the general application.

To explore matches of your interests with those of BISI faculty, see the BISI website,

For more admissions information or to apply to the program, please visit our Graduate School website: 

Application Deadlines

Type of Applicant Fall
Domestic Applicants:
US Citizens and Permanent Residents
1 Dec
International Applicants:
F (student) or J (exchange visitor) visas
A,E,G,H,I and L visas and immigrants.
1 Dec

Other Deadlines: Please visit the program website at

Degree Requirements

Master of Science (M.S.)

Total CreditsCore Requirements

Thesis option: 30 credits

Non-thesis option: 30 credits

Thesis students must complete six credits of BISI 799: Master's Thesis Research. Non-thesis students must complete a scholarly paper. All students must complete the following requirements:

• Of the 30 credits required, 18 credits must be at the 600-level or above including BISI 701: Science Teaching & Professional Development for Biology (1 credit) and BISI 688B: Bioethics (1 credit).

• Of the 30 credits required, 16 credits must be in Biological Sciences with at least three courses in a single area of specialization.


*Students cannot be formally admitted into the M.S. program, but can earn the M.S. along the way to the Ph.D.


Doctor of Philosophy (Ph.D.)

Core Requirements

All students are required to complete a minimum of 12 credits in BISI 899: Doctoral Dissertation Research, in addition to three credits of graduate seminar courses, nine credits of advanced coursework, a course in Bioethics, and BISI 701: Science Teaching & Professional Development for Biology. Students have four concentration options with additional requirements in each. 

Behavior, Ecology, Evolution, and Systematics (BEES) concentration:


BEES 608A: Seminar in Behavior, Ecology, Evolution and Systematics (1 credit)

Approved Statistics course or quantitative course (3 credits)

Computational Biology, Bioinformatics, and Genomics (CBBG) concentration:


CBMG 688Y: Bioinformatics and Genomics (2 credits)

CBMG 688P: Programming for Biology (2 credits)

CBMG 699D: Bioinformatics and Computation Biology Seminar Series (2 credits)

CBMG 688B: Bioethics (1 credit)

Electives (5 credits)

Molecular and Cell Biology (MOCB) concentration:


CBMG 688D: Cell Biology I: Structure and Function (2 credits) 

BCHM 661: Nucleic Acids (2 credits)

CBMG 688F: Gene Expression (2 credits)

CBMG 688I: Genetic Analysis (2 credits)

CBMG 688B: Bioethics (2 credits)

Electives (8 credits)

Physiological Systems (PSYS) concentration:


Bioethics course (3 credits)

Graduate Seminar course (1-3 credits) 

Three core graduate level courses (2-4 credits each)

Statistics or quantitative course (3 credits)


The Ph.D. program in Biological Sciences is an interdepartmental umbrella research program that provides opportunities for students to develop scholarly, innovative, and independent work. Courses are designed to strengthen and complement the student's research. An advisory committee helps guide each student in selecting classes and other learning experiences. Students are encouraged to present their research at national and international meetings and to publish in peer reviewed journals. Seminar series featuring prominent scientists expose students to exciting topics and help students develop collaborative contacts. During the course of their studies, each student must pass a qualifying exam, complete and defend an original dissertation, and present that dissertation work in a seminar.


Facilities and Special Resources

The campus and local area provide students access to a vast array of instrumentation, equipment, facilities, and technologies to advance biological research. As examples, the college has state of the art facilities for research in all aspects of cell and molecular biology including cell and organism culturing, protein and nucleic acid analyses, peptide sequencing, oligonucleotide synthesis and sequencing, fluorescence, confocal microscopy, scanning and transmission electron microscopy, computer graphics for molecular modeling, NMR, mass-spectroscopy, and X-ray diffraction. Students have access to a laboratory for evolutionary molecular sequence analysis; gas source stable isotope mass spectrophotometer; bioacoustic lab; flume lab; GIS (graphic information systems) lab; and high-speed network access to a wide range of desktop and super-computing facilities. Greenhouses and animal care facilities are available.

We also have several state-of the-art shared instrumentation laboratories. Two center around biological imaging for both electron and light microscopy, including a field-emission scanner and an image reconstruction/deconvolution microscope. Another shared laboratory augments existing sequencing facilities on campus, enabling large-scale processing and sequencing of nucleic acids, with multiple robotic sequenators and real time PCR. Other core facilities provide instrumentation for fluorescence-activated cell sorting (FACS), NMR, mass spectrometry, and microarray technology. Equipment and analytical instruments are available in both faculty and core laboratories for the maintenance of animal and plant tissue cultures, for the production of monoclonal antibodies, for the synthesis and micro-analysis of proteins, for large-scale fermentation and cultivation of microorganisms, and for computer assisted molecular modeling. Support staffing in shared instrumentation facilities is provided by the college, and maintenance costs have been subsidized by the college, thereby providing even occasional users with appropriate training and access, and simultaneously keeping instrument use costs low. This strategy provides exceptional opportunities for research and training, and enables graduate students to perform experiments with instrumentation that is at the leading edge of biological technology.

Students have access to the Smithsonian National Museum and USDA collections of living and preserved organisms.

Library Facilities: The library facilities on campus, as well as their online accessibility, are outstanding. In addition, there are libraries in the local area with specialized collections. The most important are the National Agricultural Library, the Library of Congress, the National Library of Medicine, and the Smithsonian Institution Library. Thus, the University of Maryland’s region contains perhaps the most comprehensive collections of books and journals in the world.

Financial Assistance

Students are supported through fellowships, research assistantships, and/or teaching assistantships. Each type of funding provides a stipend, tuition remission, and access to health and dental insurance and a prescription drug plan. Historically, all students have been supported throughout their graduate careers.

Fellowships are offered on a competitive basis.  There are no separate financial disclosure forms to fill out as part of the graduate application process.

Teaching assistantships require students to assist a faculty member in teaching a course or lab section(s). Benefits of teaching assistantships include building communication and organizational skills as well as resume enhancement for academic, government, or private sector jobs. It is also delightfully rewarding to explain concepts to students and then witness their excitement as ideas "click" and their questions are resolved.

Contact Statement

Students are strongly encouraged to communicate directly with faculty in the area of their interest. Additional general information may be obtained by emailing or by calling the Biological Sciences Graduate Office at 301-405-6905.

Please visit the Biological Sciences Graduate Program website, featuring a search engine to match research interests with faculty and links to all Concentration Areas:

The Graduate School provides detailed information on International Student Admission here: and additional information for international graduate students is available on the International Students and Scholars website at .



Gwen Warman, Program Coordinator

2101 Bioscience Research Building

University of Maryland

College Park MD, 20740

Telephone: 301-405-6905


Associated Information











Cell Biology, Developmental Biology, Molecular Biology, Genetics, Molecular Genetics, Genomics, Microbiology, Immunology, Microbial Pathogenesis, Virology, Plant Biology, Molecular Evolution,Apoptosis; Behavior; Bioenergetics; Biophysics; Cell Biology; Cell Physiology; Ecology; Estuarine and Marine Biology; Ethology; Genetics; Mitochondria; Membrane; Membrane Biophysics; Membrane Channels; Molecular Biology; Neurobiology; Physiology; Systematics; Evolutionary Biology; Evolutionary Developmental Biology; Conservation; Plant Ecology; Population Ecology; Evolution; Population Genetics; Speciation; Molecular Ecology; Restoration Ecology; Stream Ecology; Transport; Human Evolution.


Biochemistry (BCHM)

Biophysics (BIPH)

Chemistry (CHEM)

Entomology (ENTM)

Marine, Estuarine, and Environmental Sciences (MEES)

Neurosciences and Cognitive Science (NACS)